THE WORLDWIDE FRUITS
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    • ORDER ARECALES >
      • FAMILY ARECACEAE >
        • SUBFAMILY ARECOIDEAE >
          • Tribe Areceae >
            • Subtribe Arecinae >
              • Genus Areca >
                • Areca catechu - Petel nut palm
                • For Copy
            • Subtribe Euterpeinae >
              • Genus Euterpe >
                • Euterpe oleracea
                • Euterpe precatoria
            • Subtribe Iquanurinae
            • Subtribe Ptychospermatinae
            • Subtribe Roystoneinae
          • Tribe Caryoteae >
            • Genus Arenga >
              • Arenga pinnata - Sugar palm
          • Tribe Cocoeae >
            • Genus Cocos >
              • Cocos nucifera (Coconut palm)
            • Genus Elaeis >
              • Elaeis quineensis - Oil palm tree
        • SUBFAMILY CALAMOIDEAE >
          • Tribe Calameae >
            • Genus Salacca >
              • Salacca wallichiana - Rakum palm
              • Salacca zalacca - Salak
        • SUBFAMILY CEROXYLOIDEAE >
          • Genus Phytelephas >
            • Phytelephas aequatorialis
            • Phytelephas macrocarpa
        • SUBFAMILY CORYPHOIDEAE >
          • Tribe Borasseae >
            • Genus Borassus >
              • Borassus flabellifer (Toddy palm)
            • Genus Lodoicea >
              • Lodoicea maldivica (Double coconut)
          • Tribe Phoeniceae >
            • Genus Phoenix >
              • Phoenix dactylifera (Date palm)
              • Phoenix reclinata (Senrgal date palm)
              • Phoenix sylvestris (Indian date palm)
        • SUBFAMILY NYPOIDEAE >
          • Genus Nypa >
            • Nypa fruticans (Nipa palm)
    • ORDER BRASSICALES >
      • FAMILY CAPPARACEAE >
        • Genus Capparis >
          • Capparis spinosa - Caper berry
        • Genus Crateva >
          • Crateva adansonii
          • Crateva religiosa - Garlic pear
      • FAMILY CARICACEAE >
        • Genus Carica >
          • Carica papaya - Papaya
        • Genus Jacaratia
        • Genus Vasconcellea >
          • Vasconcellea candicans - Mito tree
          • Vasconcellea goudotiana
          • Vasconcellea monoica
          • Vasconcellea pubescens - Mountain papaya
          • Vasconcellea querciflora
          • Vasconcellea x heilbornii - Babaco tree
    • ORDER CARYPHYLLALES >
      • FAMILY CACTACEAE >
        • SUBFAMILY CACTOIDEAE >
          • Tribe Cacteae
          • Tribe Cereeae >
            • Genus Cereus >
              • Cereus hildmannianus
              • Cereus jamacaru
              • Cereus repandus
          • Tribe Helocereeae >
            • Genus Hylocereus >
              • Hylocereus costaricensis (Red-fleshed Pitahaya)
              • Hylocereus indatus (Pitahaya or Dragon fruit)
              • Hylocereus megalanthus (Yellow pitahaya)
        • SUBFAMILY OPUNTIEAE >
          • Tribe Opuntieae >
            • Genus Opuntia >
              • Opuntia ficus-indica (Prickly pear)
              • Opuntia matudae (Xoconostle cactus)
        • SUBFAMILY PERESKIOIDEAE >
          • Genus Pereskia >
            • Pereskia aculeata
            • Pereskia bleo - Rose cactus
    • ORDER CUCURBITALES >
      • FAMILY CUCURBITACEAE >
        • SUBFAMILY CUCURBITOIDEAE >
          • Tribe Benincaseae >
            • Subtribe Benincasinae >
              • Genus Acanthosicyos >
                • Acanthosicyos horridus - Nara melon
                • Acathosicyos naudinianus - Gemsbok cucumber
              • Genus Benincasa >
                • Benincasa hispida - Winter melon
              • Genus Coccinia >
                • Coccinia grandis - Ivy gourd
              • Genus Cucumis >
                • Cucumis anguria - Maroon cucumber
                • Cucumis dipsaceus - Wild Spiny Cucumber
                • Cucumis melo - Muskmelon >
                  • Cantaloupe
                • Cucumis humofructus - Aadvark cucumber
                • Cucumis metuliferus - Horned melon
                • Cucumis myriocarpus - Gooseberry cucumber
                • Cucumis prophetarum - Paddy melon
                • Cucumis sativus (Cucumber) >
                  • Lemon Cucumber
              • Genus Citrullus >
                • Citrullus caffer - Citron melon
                • Citrullus colocynthis - Desert gourd
                • Citrullus ecirrhosus - Tsamma melon
                • Citrullus lanatus - Watermelon
              • Genus Lagenaria >
                • Lagennaria siceraria - Bottle gourd
            • Subtribe Luffinae >
              • Subtribe Cucumerinae
          • Tribe Cucurbiteae >
            • Genus Abobra >
              • Abobra tenuifolia - Cranberry gourd
            • Genus Cucurbita >
              • Cucurbita argyrosperma - Cushaw pumpkin
              • Cucurbita ficifolia - Figleaf gourd
              • Cucurbita foetidissima - Buffalo gourd
              • Cucurbita maxima - Giant pumpkin
              • Cucurbita moschata - Butternut squash
              • Cucurbita okeechobeensis
              • Cucurbita pepo - Field pumpkin
            • Genus Sicana >
              • New Page
              • Acorn Squash
          • Tribe Joliffieae >
            • Genus Siraitia >
              • Siraitia grosvenorii (Luo han guo or Monk fruit)
            • Genus Telfairia
          • Tribe Momordiceae >
            • Genus Momordica >
              • Momordica charantia - Bitter melon
              • Momordica cochinchinensis - Gac
              • Momordica dioica - Spine Gourd
          • Tribe Sicyoeae >
            • Genus Cyclanthera >
              • Cyclanthera explodens
              • Cyclanthera pedata - Achocha gourd
            • Genus Frantzia
            • Genus Hodgsonia
            • Genus Luffa >
              • Luffa accutangula (Angled luffa)
              • Luffa Aegyptiaca (Smooth luffa)
            • Genus Marah
            • Genus Sechium - Chayota genus >
              • Sechium edule - Chayote
            • Genus Trichosanthes >
              • Trichosanthes cucumerina - Snake gourd
          • Tribe Siraitieae >
            • Genus Siraitia >
              • Siraitia grosvenorii - Monk fruit
    • ORDER ERICALES >
      • FAMILY ACTINIDIACEAE >
        • Genus Actinidia >
          • Actinidia arguta - Hardy kiwifruit
          • Actinidia chinensis - Golden kiwi)fruit
          • Actinidia diliciosa - Kiwi fruit
          • Actinidia kolomikta - Arctic kiwifruit
          • Actinidia melanandra - Red kiwifruit
          • Actinidia polygama - Silver vine
      • FAMILY EBENACEAE >
        • Genus Diospyros >
          • Diospyros blancoi - Velvet persimmon
          • Diospyros decandra - Gold apple
          • Diospyros kaki - Asian Persimmon
          • Diospyros lotus - Lilac persimmon
          • Diospyros nigra - Black sapote
          • Diospyros taxana - Texas persimmon
          • Diospyros virginiana (American Persimmon)
      • FAMILY ERICACEAE >
        • SUBFAMILY ARBUTOIDEAE >
          • Genus Arbutus >
            • Arbutus andrachne - Greek strawberry tree
            • Arbutus azizonica
            • Arbutus unedo - Strawberry tree
            • Arbutus xalapensis
          • Genus Artostaphylos
          • Genus Xylococcus >
            • Xylococcus bicolor - Mission mananita
        • SUBFAMILY STYPHELIOIDEAE >
          • Genus Leptecophylla >
            • Leptecophylla juniperina
            • Leptecophylla tameiameiae
          • Genus Styphelia
        • SUBFAMILY VACCINOIDEAE >
          • Tribe Gaultherieae >
            • Genus Gaultheria
          • Tribe Vaccinieae >
            • Genus Gaylussacia >
              • Gaylussacia baccata - Black Huckleberry
              • Gaylussacia branchycera (Box Huckleberry)
              • Gaylussacia dumosa (Dwarf Huckleberry)
              • Gaylussacia frondosa (Blue Huckleberry)
            • Genus Vaccinium >
              • Bilberry or Whortleberry
              • Blueberry
              • Cranberry
              • Lingonberry or Cowberry
              • Huckleberry
      • FAMILY LECYTHIDACEAE >
        • Genus Bertholletia >
          • Bertholletia excelsa - Brazil nut
        • Genus Couroupita >
          • Couroupita guianensis - Cannonball tree
        • Genus Gustavia >
          • Gustavia superba
        • Genus Lecythis >
          • Lecythis pisonic
          • Lecythis zabucajo
      • FAMILY SAPOTACEAE >
        • SUBFAMILY CHRYSOPHYLLOIDEAE >
          • Genus Chrysophyllum >
            • Chrysophyllum albidum - African star apple
            • Chrysophyllum cainito - Milk fruit
            • Chrysophyllum oliviforme - Wild Star apple
            • Chrysophyllum roxburghii
            • Chrysophyllum magalismontanum
          • Genus Englerophytum >
            • Englerrophytum magalismontanum
            • Englerophytum natalense
          • Genus Synsepalum >
            • Synsepalum dulcificum (Miracle fruit)
          • Genus Pouteria >
            • Pouteria australis
            • Pouteria campechiana - Cannistel
            • Pouteria caimito - Abiu tree
            • Pouteria eerwark
            • Pouteria lucuma - Lucuma
            • Pouteria multiflora - Broad - leaved lucuma
            • Pouteria sandwicensis
            • Pouteria sapota - Mamey sapote
            • Pouteria viridis - Green sapote
        • SUBFAMILY SAPODOIDEAE >
          • Tribe Sapoteae >
            • Genus Manilkara >
              • Manilkara bidentata - Cow tree
              • Manilkara chicle (Chicle)
              • Manilkara hexandra
              • Manilkara huberi
              • Manilkara zopota (Sapodilla)
            • Genus Mimusops >
              • Mimusops caffra
              • Mimusops elengi
              • Mimusops zeyheri
          • Tribe Sideroxyleae >
            • Genus Argania >
              • Argania spinosa
    • ORDER FABALES >
      • FAMILY FABACEAE >
        • SUBFAMILY CAESALPINIOIDEAE >
          • Genus Pithecellobium >
            • Pithecellobium dulce
        • SUBFAMILY DETARIOIDEAE >
          • Genus Tamarindus - Tamarind genus >
            • Tamarindus indica - Tamarind
        • SUBFAMILY DIALIOIDEAE >
          • Genus Dialium >
            • Dialium cochinchinense
            • Dialium guineense - Black velvet tamarind
            • Dialium indum - Tamarin plum
        • SUBFAMILY FABOIDEAE >
          • Tribe Dalbergieae >
            • Genus Arachis - Peanut genus >
              • Arachis hypogaea - Peanut
          • Tribe Fabeae >
            • Genus Lathyrus - Peavine genus
            • Genus Pisum >
              • Pisum sativum - Pea
            • Genus Vicia >
              • Vicia faba - Broad bean
          • Tribe Phaseoleae >
            • Subtribe Cajaninae >
              • Genus Cajanus - Pigeon pea genus >
                • Cajanus cajan - Pigeon pea
            • Subtribe Glycininae >
              • Genus Glycine >
                • Glycine max
            • Subtribe Phaseolinae >
              • Genus Psophocarpus - Winged bean genus >
                • Psophocarpus tetragonolobus - Winged bean
    • ORDER LAMIALES >
      • FAMILY OLEACEAE >
        • Tribe Oleeae >
          • Genus Olea >
            • Olea europaea - Olive
      • FAMILY PEDALIACEAE >
        • Genus Sesamum >
          • Sesamum indicum - Sesame
          • Genus Couroupita
    • ORDER LAURALES >
      • FAMILY LAURACEAE >
        • Genus Persea >
          • Persea americana (Avocado)
    • ORDER MAGNOLIALES >
      • FAMILY ANNONACEAE >
        • Genus Annona >
          • Annona x atemoya - Atenmoya
          • Annona cherimola - Cherimoya
          • Annona chrysophylia - Graines
          • Annona crassiflora - Marrolo
          • Annona diversiflora - Ilama
          • Annona glabra - Pond apple
          • Annona montana - Mountain soursop
          • Annona muricata - Soursop
          • Annona purpurea - Soncoya
          • Annona reticulata - Custard apple
          • Annona sebegalensis - African custard apple
          • Annona squamosa - Sweetsop
        • Genus Anonidium >
          • Anonidium mannii - Junglesop
        • Genus Asimina >
          • Asimina triloba - Common pawpaw
        • Genus Monodora >
          • Monodora myristica - Calabash nutmeg
        • Genus Rollinia >
          • Rollinia deliciosa - Wild sugar apple
        • Genus Uvaria >
          • Uvaria chamae
          • Uvaria ovata
          • Uvaria rufa
      • FAMILY MYRRISTICACEAE >
        • Genus Myristica >
          • Myristica fragrans - True nutmeg
          • Myristica malabarica - Bombay nutmeg
        • Genus Pycnanthus
    • ORDER MALPIGHIALES >
      • FAMILY CALOPHYLLACEAE >
        • Genus Mammea >
          • Mammea africana - African mamey apple
          • Mammea americana - Mamey apple
      • FAMILY CLUSIACEAE >
        • SUBFAMILY CLUSIOIDEAE >
          • Tribe Garcinieae >
            • Genus Garcinia >
              • Garcinia dulcis - Mundu tree
              • Garcinia humilis - Achacha tree
              • Garcinia gardneriana - Bacupari
              • Garcinia indica - Kokum tree
              • Garcinia intermedia - Lemon drop mangosteen
              • Garcinia mangostana - Mangosteen
          • Tribe Platonieae >
            • Genus Platonia - Bacury genus >
              • Platonia insignis - Bacuri tree
          • Tribe Plukenetieae
        • SUBFAMILY HYPERICOIDEAE >
          • Tribe Calophylleae >
            • Genus Mammea
      • FAMILY EUPHORBIACEAE >
        • SUBFAMILY ACALYPHOIDEAE >
          • Tribe Acalypheae >
            • Genus Ricinus >
              • Ricinus communis
          • Tribe Plukenetieae >
            • Genus Plukenetia >
              • Plukenetia conophora
              • Plukenetia volubitis
        • SUBFAMILY CROTONOIDEAE >
          • Tribe Aleuritideae >
            • Genus Aleurites
          • Tribe Manitroteae >
            • Genus Manihot
          • Tribe Micrandreae >
            • Genus Schinziophyton
      • FAMILY MALPIGHIACEAE >
        • Genus Bunchosia >
          • Bunchosia argentea
          • Bunchosia armeniaca >
            • FAMILY GOMORTEGACEAE >
              • Genus Gomortega >
                • Gomortega keule
        • Genus Byrsonima >
          • Byrsonima crassifolia - Nance fruit
        • Genus Malpighia >
          • Malpighia emarginata (West Indian cherry)
      • FAMILY PHYLLANTHACEAE >
        • SUBFAMILY ANTIDESMATOIDEAE >
          • Tribe Antidesmeae >
            • Genus Antidesma >
              • Antidesma bunius
          • Tribe Scepinae >
            • Genus Baccaurea
        • SUBFAMILY PHYLLANTHOIDEAE >
          • Tribe Phyllantheae >
            • Genus Phyllanthus >
              • Phyllanthus acidus (Star gooseberry)
              • Phyllanthus emblica (Indian gooseberry)
    • ORDER MALVALES >
      • FAMILY BIXACEAE >
        • Genus Bixa >
          • Bixa orellana
      • FAMILY MALVACEAE >
        • SUBFAMILY BOMBACOIDEAE >
          • Genus Adansona - Baobab genus >
            • Adansonia digitata - Baobab
            • Adansonia grandidieri - Giant baobab
            • Andansonia gregorii - Australian baobab
          • Genus Quararibea >
            • Quararibea cordata - South American Sapote
        • SUBFAMILY BYTTNEROIDEAE >
          • Genus Theobroma >
            • Theobroma bicolor (Mocambo)
            • Theobroma cacao (Cacao tree)
            • Theobroma granddiflorum - Cupuacu tree
        • SUBFAMILY HELICTEROIDEAE >
          • Tribe Durionneae >
            • Genus Durio L. >
              • Durio dulcis (Red Durian)
              • Durio kutejensis - Durian pulu
              • Durio zibethinus (Durian)
              • Durio graveolens - Red fleshed durian
        • SUBFAMILY MALVOIDEAE >
          • Tribe Hibisceae >
            • Genus Abelmoschus >
              • Abelmoschus caillei - West African okra
              • Abelmoschus esculentus - Okra
        • SUBFAMILY STERCULIOIDEAE >
          • Genus Cola >
            • Cola acuminata
            • Cola nilida
          • Genus Sterculia
      • FAMILY MUNTINGIACEAE >
        • Genus Muntingia >
          • Muntingia calabura (Jamaica cherry)
    • ORDER MYRTALES >
      • FAMILY LYTHRACEAE >
        • SUBFAMILY PUNICOIDEAE >
          • Genus Punica >
            • Runica granatum (Pomegranate)
            • Punica protopunica (Socotran pomegranate)
        • SUBFAMILY SONNERATIOIDEAE >
          • Genus Sonneratia >
            • Sonneratia alba
            • Sonneratia caseolaris
            • Sonneratia hainanensis
            • Sonneratia ovata
        • SUBFAMILY TRAPOIDEAE >
          • Genus Trapa >
            • Trapa natan - Water caltrop
      • FAMILY MYRTACEAE >
        • SUBFAMILY MYRTOIDEAE >
          • Tribe Myrteae >
            • Genus Acca >
              • Acca sellowiana
            • Genus Amomyrtus
            • Genus Austromyrtus
            • Genus Eugenia
            • Genus Hexachlamys
            • Genus Luma >
              • Luma apiculata
              • Luma chequen
            • Genus Myrtus
            • Genus Plinia
            • Genus Psidium >
              • Psidium cattleyanum (Cherry Guava)
              • Psidium guajava (Guava)
            • Genus Rhodomyrtus
            • Genus Ugni
          • Tribe Syzygieae >
            • Genus Syzygium >
              • Syzygium jambos
              • Syzygium oleosum
              • Syzygium paniculatum
              • Syzygium papyraceum
              • Syzygium samarangense (Java apple)
              • Syzygium smithii
              • New Page
    • ORDER ROSALES >
      • FAMILY CANNABACEAE >
        • Genus Celtis >
          • Celtis occidentalis
          • Celtis sinensis
      • FAMILY ELAEAGNACEAE >
        • Genus Elaeagnus >
          • Elaegnus angustifola - Russian olive
          • Elaeagnus multiflora - Cherry silverberry
        • Genus Hippophae >
          • Hippophae rhamnoides - Sea buckthorn
        • Genus Shepherdia >
          • Shepherdia argentea - Silver buffaloberry
          • Shepherdia canadensis - Canadian buffalberry
      • FAMILY MORACEAE >
        • Tribe Artocarpeae >
          • Genus Artocarpus >
            • Artocarpus altilis (Breadfruit)
            • Artocarpus camansi - Breadnut
            • Artocarpus integer (Cempedak)
            • Artocarpus heterophyllus (Jackfruit)
          • Genus Treculia >
            • Treculia africana (African breadfruit)
        • Tribe Dorstenieae >
          • Genus Brosimum - Breadnut genus >
            • Brosimum alicastrum - Breadnut
        • Tribe Durioneae >
          • Genus Durio >
            • Durio zibethinus (Durian)
        • Tribe Ficeae >
          • Genus Ficus >
            • Ficus carica - Common fig
            • Ficus coronata - Creek sandpeper fig
            • Ficus platypoda - Desert fig
            • Ficus racemosa - Cluster fig
        • Tribe Moceae >
          • Genus Maclura >
            • Maclura cochinchinensis - Cockpur thorn
            • Maclura pomifera - Osage orange
          • Genus Morus >
            • Morus nigra - Black mulberry
            • Morus rubra - Red mulberry
      • FAMILY RHAMNACEAE >
        • Genus Ziziphus >
          • Ziziphus jujuba
          • Ziziphus lotus
          • Ziziphus mauritiana
          • Ziziphus mistol
          • Ziziphus mucronata
          • Ziziphus nummularia
      • FAMILY ROSACEAE >
        • SUBFAMILY AMYGDALOIDEAE >
          • Tribe Amygdaleae >
            • Genus Prunus >
              • Subgenus Amygdalus - Almond and Peach >
                • Prunus dulcis - Almond
                • Prunus persica - Peach
              • Subgenus Cerasus - Cherry tree >
                • Prunus avium (Sweet cherry)
                • Prunus cerasus (Sourc cherry)
              • Subgenus Prunus - Apricot and Plum >
                • Section Armeniaca - Apricot trees
                • Plum trees
              • Subgenus laurocerasus - Cherry laurel
              • Subgenus Lithocerasus
              • Subgenus Padus - Bird cherry
          • Tribe Maleae >
            • Subtribe Malinae >
              • Genus Cydonia >
                • Cydonia oblonga (Quince)
              • Genus Eriobotrya >
                • Eriobotrya japonica (Loquat)
              • Genus Malus >
                • Malus domestica - Apple tree
                • Malus sylvestris - European Wild apple
                • Malus pumila (Apple tree)
              • Genus Pyrus >
                • Pyrus communis - European pear
                • Pyrus pyrifolia - Asian pear
                • Pyrus x bretschneideri - Chinese white pear
              • Genus Sorbus >
                • Sorbus subgenus Aria
                • Sorbus subgenus Chamaemespilus
                • Sorbus subgenus Cormus
                • Sorbus subgenus Micromeles
                • Sorbus subgenus Sorbus
                • Sorbus subgenus Torminaria
        • SUBFAMILY ROSOIDEAE >
          • Tribe Rubeae >
            • Genus Prunus
            • Genus Rubus >
              • Rubus fruticosus (Blackberry)
              • Rubus x loganobaccus (Loganberry)
            • Genus Eriobotrya
          • Tribe Potentilleae >
            • Genus Fragaria >
              • Fragaria x ananassa (Straw berry)
              • Fragaria virginiana (Virgina strawberry)
              • Fragaria moschata - Musk strawberry
    • ORDER SAPINDALES >
      • FAMILY ANACARDIACEAE >
        • SUBFAMILY ANACARDIOIDEAE >
          • Genus Anacardium >
            • Anacardium occidentale (Cashew tree)
            • Anacardium othonianum (Little cashew)
          • Genus Bouea
          • Genus Mangifera - Mango genus >
            • Mangifera indica - Mango tree
            • Mangifera odorata - Saipan Mango
            • Mangifera siamensis - Thai Mango
            • Mangifera sylvatica - Nepal Mango
          • Genus Rhus >
            • Rhus ovata - Sugar sumac
            • Rhus typhina - Staghorn sumac
        • SUBFAMILY SPONDIOIDEAE >
          • Genus Dracontomelon >
            • Dracontomelon duperreanum (Vietnamese: Sau)
          • Genus Spondias >
            • Spondias dulcis (Golden apple)
            • Spondias mombin (Yellow mommbin)
            • Spondias purpurea - Purple mombin
            • Spondias tuberosa (Brazil plum)
      • FAMILY BURSERACEAE >
        • Genus Canarium >
          • Canarium album
          • Canarium luzonicum
          • Canarium ovatum
          • Canarium pseudodecumanum
        • Genus Dacryodes >
          • Dacryodes edulis - Safou tree
      • FAMILY MELIACEAE >
        • Genus Lansium >
          • Lansium parasiticum - Langsat tree
        • Genus Sandoricum >
          • Sandoricum koetiape - Santol fruit
      • FAMILY RUTACEAE >
        • SUBFAMILY AURANTIOIDEAE >
          • Tribe Citreae >
            • Subtribe Balamocitrinae >
              • Genus Aegle - Bael genus
            • Subtribe Citrinae >
              • Genus Citrus >
                • Important Species >
                  • Citrus maxima (Pomelo)
                  • Citrus medica (Citron)
                  • Citrus reticulata - Mandarin
                  • Citrus tangerina (Tangerine)
                • Important Hybrids >
                  • Citrus x sinensis (Sweet orange)
                  • Citrus reticula x sinensis (King orange)
              • Genus Fortunella
            • FAMILY VITACEAE >
              • Genus Cayratia >
                • Cayralia trifola (Vietnamese wild grape)
              • Genus Vitis >
                • Vitis labrusca (Fox grape vine)
                • Vitis vinifera (Common grape vine)
            • Subtribe Triphasiinae
          • Tribe Clauseneae >
            • Genus Glycosmis
        • SUBFAMILY TODDALIOIDEAE >
          • Genus Casimiroa >
            • Casimiroa edulis - White sapote
      • FAMILY SAPINDACEAE >
        • SUBFAMILY HIPPOCASTANOIDEAE >
          • Genus Aesculus - Horse chestnut genus >
            • Aesculus hippocastanum - Horse chestnut
        • SUBFAMILY SAPINDOIDEAE >
          • Genus Blighia >
            • Blighia sapida (Ackee)
          • Genus Diploglottis >
            • Diploglottis australis - Australian tamarin
          • Genus Dimocarpus >
            • Dimocarpus longan (Longan tree)
          • Genus Litchi >
            • Litchi chinensis (Lychee)
          • Genus Melicoccus >
            • Melicoccus bijugatus - Spanish lime
          • Genus Nephelium >
            • Nephelium hypoleucum (Korlan tree)
            • Nephelium malabile (Pulasan tree)
            • Nephelium lappaceum (Rambutan)
            • Nephelium xerospermoides (Hairless Rambutan)
          • Genus Paullinia >
            • Paullinia cupana - Guarana tree
          • Genus Talisia >
            • Talisia esculenta
        • SUBFAMILY XANTHOCEROIDEAE >
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  • ABOUT FRUITS
  • Types of Fruits

Genus Solanum

Solanum lycopersicum
​Tomato plant

Edited by Ho Dinh Hai
Long An - Vietnam
​Updated: 03/03/2017

1-Introduction to the Species Solanum lycopersicum L. - Tomato plant

               1.1- Scientific classification
                Kingdom:         Plantae
               (unranked):       Angiosperms
               (unranked):       Eudicots
               (unranked):       Asterids
                Order:              Solanales
                Family:             Solanaceae
                Subfamily:       Solanoideae
                Tribe:               Solaneae
                Genus:             Solanum L.
                Species:           Solanum lycopersicum L.
                1.2- Species Solanum lycopersicum L. - Tomato plant
                + Overview
             The Tomato is the edible fruit of Solanum lycopersicum L, commonly known as a tomato plant, which belongs to the  nightshade  family, Solanaceae.
           The species originated in Central and South America. The Nahuatl (Aztec language) word tomatl gave rise to the Spanish word "tomate", from which the English word tomato originates.
             Its use as a food originated in Mexico, and spread throughout the world following the Spanish colonization of the Americas. Tomato is consumed in diverse ways, including raw, as an ingredient in many dishes, sauces, salads, and drinks. While tomatoes are  botanically  berry-type fruits, they are considered culinary vegetables, being ingredients of savory meals.
               Synonyms:
              1- Lycopersicon lycopersicum (L.) H. Karst.
              2- Lycopersicon esculentum Mill.
               + Etymology
               The word "tomato" comes from the Spanish tomate, which in turn comes from the Nahuatl word tomatl. It first appeared in print in 1595.
            The native Mexican tomatillo is tomate (in Nahuatl: tomātl ), meaning "fat water" or "fat thing"). When Aztecs started to cultivate the Andean fruit, bigger and red, they called the new species xitomatl (or jitomates), ("plump thing with navel" or "fat water with navel"). After their conquest of Tenochtitlan, Spaniards exported tomatoes (jitomates) to the rest of the world with the name tomate, so numerous languages use forms of the word "tomato" (tomate) to refer to the red tomato instead of the green tomatillo. It first appeared in print in 1595. The scientific species epithet lycopersicum means "wolf peach", and comes from German  werewolf myths. These legends said that deadly nightshade was used by witches and sorcerers in potions to transform themselves into werewolves, so the tomato's similar, but much larger, fruit was called the "wolf peach" when it arrived in Europe.

2- Characteristics of the Species Solanum lycopersicum L . - Tomato plant

​                 2.1- Description
                  + The plants
                Tomato plants are vines, initially decumbent, typically growing 180 cm (6 ft) or more above the ground if supported, although erect bush varieties have been bred, generally 100 cm (3 ft) tall or shorter. Indeterminate types are "tender" perennials, dying annually in temperate climates (they are originally native to tropical highlands), although they can live up to three years in a greenhouse in some cases. Determinate types are annual in all climates.
                Tomato plants are dicots, and grow as a series of branching stems, with a terminal bud at the tip that does the actual growing. When that tip eventually stops growing, whether because of pruning or flowering, lateral buds take over and grow into other, fully functional, vines.
The plants typically grow to 1-3 meters (3-10 ft) in height and have a weak stem that often sprawls over the ground and vines over other plants. It is a perennial in its native habitat, and grown as an annual in temperate climates.
                + The leaves
           Most tomato plants have compound leaves, and are called regular leaf (RL) plants, but some cultivars have simple leaves known as potato leaf (PL) style because of their resemblance to that particular relative. Of RL plants, there are variations, such as rugose leaves, which are deeply grooved, and variegated, angora leaves, which have additional colors where a genetic mutation causes chlorophyll to be excluded from some portions of the leaves.
            The leaves are 10-25 cm (4-10 in) long, odd pinnate, with five to 9 leaflets on petioles, each leaflet up to 8 cm (3 in) long, with a serrated margin; both the stem and leaves are densely glandular-hairy.
                + The flowers
             Their flowers, appearing on the apical meristem, have the anthers fused along the edges, forming a column surrounding the pistil's style. Flowers in domestic cultivars can be self-fertilizing. The flowers are 1-2 cm (0.4-0.8 in) across, yellow, with five pointed lobes on the corolla; they are borne in a cyme of three to 12 together.
                 + The fruits
               Tomato fruit is classified as a berry. As a true fruit, it develops from the ovary of the plant after fertilization, its flesh comprising the pericarp walls. The fruit contains hollow spaces full of seeds and moisture, called locular cavities. These vary, among cultivated species, according to type. Some smaller varieties have two cavities, globe-shaped varieties typically have three to five, beefsteak tomatoes have a great number of smaller cavities, while paste tomatoes have very few, very small cavities.
                 + The seeds
                 Each fruit contains at least two cells of small seeds surrounded by jellylike pulp.
                 For propagation, the seeds need to come from a mature fruit, and be dried or fermented before germination.
Picture
Many types of Tomatoes
                 2.2- Origin and Distribution
                 + Origin:
                The tomato is native to western South America and Central America. Native versions were small, like cherry tomatoes, and most likely yellow rather than red.
                 + Distribution:
                Tomatoes originated in South America but are now found all over the world.
                The tomato is grown worldwide for its edible fruits, with thousands of cultivars.
               In 2014, world production of tomatoes was 170.8 million tonnes, with China accounting for 31% of the total, followed by India, the United States and Turkey as the major producers.
                The major producers of Tomatoes in 2014 are: 1- China, 2- India, 3- United States, 4- Turkey, 5- Egypt, 6- Iran, 7- Italy and 8- Spain
                 2.3- Biological characteristics
                  + Note in Classification
           In 1753, Linnaeus placed the tomato in the genus Solanum (alongside the potato) as  Solanum lycopersicum. In 1768, Philip Miller moved it to its own genus, naming it  Lycopersicon esculentum. This name came into wide use, but was in breach of the plant naming rules. Although the name Lycopersicon lycopersicum may be found, it is not used because it violates the International Code of Nomenclature barring the use of tautonyms in botanical nomenclature.
              Genetic evidence has now shown that Linnaeus was correct to put the tomato in the genus Solanum, making Solanum lycopersicum  the correct name. Both names, however, will probably be found in the literature for some time. Two of the major reasons that some still consider the genera separate are the leaf structure (tomato leaves are markedly different from any other Solanum), and the biochemistry (many of the alkaloids common to other  Solanum  species are conspicuously absent in the tomato). Hybrids of tomato and diploid  potato can be created in the lab by somatic fusion, and are partially fertile, providing evidence of the close relationship between these species.
                + Fruit versus vegetable
                Botanically, a tomato is a fruit, a berry, consisting of the ovary, together with its seeds, of a flowering plant. However, the tomato has a much lower sugar content than other edible fruits, and is therefore not as sweet. Typically served as part of a salad or main course of a meal, rather than at dessert, it is, in the US, considered a "culinary vegetable". One exception is that tomatoes are treated as a fruit in home canning practices: they are acidic enough to process in a water bath rather than a pressure cooker as vegetables require. Tomatoes are not the only food source with this ambiguity: bell peppers, cucumbers, green beans, eggplants, avocados, and squashes of all kinds (such as  zucchini  and  pumpkins) are all botanically fruits, yet cooked as vegetables. This has led to legal dispute in the United States. In 1887, US tariff laws that imposed a dutyon vegetables, but not on fruits, caused the tomato's status to become a matter of legal importance. The US Supreme Court settled this controversy on 10 May 1893, by declaring that the tomato is a vegetable, based on the popular definition that classifies vegetables by use, that they are generally served with dinner and not dessert (Nix v. Hedden (149 U.S. 304)). The holding of this case applies only to the interpretation of the Tariff Act of 3 March 1883, and the court did not purport to reclassify the tomato for botanical or other purposes.
                 + Pollination
                In the wild, original state, tomatoes required cross-pollination; they were much more  self-incompatible than domestic cultivars. As a floral device to reduce selfing, the pistil of wild tomatoes extends farther out of the flower than today's cultivars. The stamens were, and remain, entirely within the closed corolla.
               As tomatoes were moved from their native areas, their traditional pollinators, (probably a species of halictid bee) did not move with them. The trait of self-fertility became an advantage, and domestic cultivars of tomato have been selected to maximize this trait.
               This is not the same as self-pollination, despite the common claim that tomatoes do so. That tomatoes pollinate themselves poorly without outside aid is clearly shown in  greenhouse  situations, where pollination must be aided by artificial wind, vibration of the plants (one brand of vibrator is a wand called an "electric bee" that is used manually), or more often today, by cultured bumblebees. The anther of a tomato flower is shaped like a hollow tube, with the pollen produced within the structure, rather than on the surface, as in most species. The pollen moves through pores in the anther, but very little pollen is shed without some kind of externally-induced motion. The best source of outside motion is a sonicating bee, such as a bumblebee, or the original wild halictid pollinator. In an outdoors setting, wind or animals usually provide sufficient motion to produce commercially viable crops.
                 + Genetic modification
              Tomatoes that have been modified using genetic engineering have been developed, and although none are commercially available now, they have been in the past. The first commercially available genetically modified food was a variety of tomato named the Flavr Savr, which was engineered to have a longer shelf life. Scientists are continuing to develop tomatoes with new traits not found in natural crops, such as increased resistance to pests or environmental stresses. Other projects aim to enrich tomatoes with substances that may offer health benefits or provide better nutrition.
                An international consortium of researchers from 10 countries, among them researchers from the Boyce Thompson Institute for Plant Research, began sequencing the tomato  genome  in 2004, and is creating a database of genomic sequences and information on the tomato and related plants. A prerelease version of the genome was made available in December 2009. The genomes of itsmitochondria and chloroplasts are also being sequenced as part of the project. The complete genome for the cultivar Heinz 1706 was published on 31 May 2012 in Nature. Since many other fruits, like strawberries, apples, melons, and bananas share the same characteristics and genes, researchers stated the published genome could help to improve food quality, food security and reduce costs of all of these fruits.
                See also: Genetically modified tomato
                 + Breeding
               The Tomato Genetic Resource Center, Germplasm Resources Information Network,  AVRDC, and numerous seed banks around the world store seed representing genetic variations of value to modern agriculture. These seed stocks are available for legitimate breeding and research efforts. While individual breeding efforts can produce useful results, the bulk of tomato breeding work is at universities and major agriculture-related corporations. These efforts have resulted in significant regionally adapted breeding lines and hybrids, such as the Mountain series from North Carolina. Corporations including Heinz, Monsanto, BHNSeed, and Bejoseed have breeding programs that attempt to improve production, size, shape, color, flavor, disease tolerance, pest tolerance, nutritional value, and numerous other traits.
                  + Plant toxicity
               Leaves, stems, and green unripe fruit of the tomato plant contain small amounts of the toxic alkaloid tomatine. They also contain small amounts of solanine, a toxic alkaloid found in potato leaves and other plants in the nightshade family. Use of tomato leaves in herbal tea has been responsible for at least one death. However, levels of tomatine in foliage and green fruit are generally too small to be dangerous unless large amounts are consumed, for example, as greens. Small amounts of tomato foliage are sometimes used for flavoring without ill effect, and the green fruit is sometimes used for cooking, particularly as fried green tomatoes. Compared to potatoes, the amount of solanine in green or ripe tomatoes is low; however, even in the case of potatoes while solanine poisoning resulting from dosages several times normal human consumption has been demonstrated, actual cases of poisoning resulting from excessive consumption of potatoes that have high concentration of solanine are rare.
                  Tomato plants can be toxic to dogs if they eat large amounts of the fruit, or chew plant material.
                  2.4- Nutrition of Tomato fruits
                  + Introduction
               A tomato is 95% water, contains 4% carbohydrates and less than 1% each of fat and protein (table). In a 100 gram amount, raw tomatoes supply 18 calories and are a moderate source of vitamin C (17% of the Daily Value), but otherwise are absent of significant nutrient content (See table).
                  + Nutritional value per 100 g (3.5 oz) of Red tomatoes, raw
                 Energy....................................................
                 Carbohydrates
                 Sugars
                 Dietary fiber
                 Fat
                 Protein
                  Vitamins................................................
                 Vitamin A equiv.
                 Beta-carotene
                 Lutein zeaxanthin
                 Thiamine (B1)
                 Niacin (B 3 )
                 Vitamin B6
                 Vitamin C
                 Vitamin E
                 Vitamin K
                  Minerals................................................
                 Magnesium
                 Manganese
                 Phosphorus
                 Potassium
                  Other constituents................................
                 Water
                 Lycopen
​74 kJ (18 kcal)
3.9 g
2.6 g
1.2 g
0.2 g
0.9 g
:
42 μg (5%)
449 μg (4%)
123 μg
0.037 mg (3%)
0.594 mg (4%)
0.08 mg (6%)
14 mg (17%)
0.54 mg (4%)
7.9 μg (8%)
:
11 mg (3%)
0.114 mg (5%)
24 mg (3%)
237 mg (5%)
:
94.5 g
2573 µg
                 Note!
                + Units: μg = micrograms • mg = milligrams * IU = International units
                + Percentages are roughly approximated using US recommendations for adults.
                 Source: USDA Nutrient Database - Link to USDA Database entr
                2.5- Health Benefits of Tomato fruits
                 + Introduction
                Tomatoes are a rich source of vitamins A and C and folic acid. Tomatoes contain a wide array of beneficial nutrients and antioxidants, including alpha-lipoic acid, lycopene, choline, folic acid, beta-carotene and lutein.
              One medium tomato (approximately 123 grams) provides 22 calories, 0 grams of fat, 5 grams of carbohydrate (including 1 gram of fiber and 3 grams of sugar) and 1 gram of protein.
              Cooking tomatoes appears to increase the availability of key nutrients such as the caroteinoids lycopene, lutein and zeaxanthin. Stewed tomatoes provide considerably more lutein and zeaxanthin than sun dried tomatoes and raw cherry tomatoes.
               Alpha-lipoic acid helps the body to convert glucose into energy. Some evidence suggests that alpha-lipoic acid can aid in blood glucose control, improve vasodilation and protect against retinopathy in people with diabetes; it may even help preserve brain and nerve tissue.
               Lycopene is the antioxidant that gives tomatoes their rich red color. Tomatoes account for 80 percent of lycopene consumption in the average diet.
              Choline is an important nutrient found in tomatoes that helps with sleep, muscle movement, learning and memory. Choline also helps to maintain the structure of cellular membranes, aids in the transmission of nerve impulses, assists in the absorption of fat and reduces chronic inflammation.
                + Possible health benefits of tomatoes
               Tomatoes are packed full of beneficial nutrients and antioxidants and are a rich source of vitamins A and C and folic acid.
              The benefits of consuming fruits and vegetables of all kinds, including tomatoes, are impressive. As the proportion of plant foods in the diet increases, the risk of heart disease, diabetes, and cancer goes down.
            High fruit and vegetable intake is also associated with healthy skin and hair, increased energy and lower weight. Increasing consumption of fruits and vegetables significantly decreases the risk of obesity and overall mortality.
                   + The 11 Health Benefits of Tomatoes
                   1)- Cancer
                 As an excellent source of vitamin C and other antioxidants, tomatoes can help combat the formation of free radicals known to cause cancer.
                   2)- Prostate Cancer
                  Lycopene has been linked with prostate cancer prevention in several studies. According to John Erdman, Ph.D., Professor Emeritus of the department of food science and human nutrition at the University of Illinois, "There's very good, strong, epidemiological support for increased consumption of tomato products and lower incidence of prostate cancer."
                  Among younger men, diets rich in beta-carotene may play a protective role against prostate cancer, according to a study conducted by the Harvard School of Public Health's Department of Nutrition.
                 3)- Colorectal Cancer
            Beta-carotene consumption has been shown to have an inverse association with the development of colon cancer in the Japanese population. High fiber intakes from fruits and vegetables are associated with a lowered risk of colorectal cancer.
               According to the American Cancer Society, some studies have shown that people who have diets rich in tomatoes may have a lower risk of certain types of cancer, especially cancers of the prostate, lung, and stomach. Further human-based research is needed to find out what role lycopene might play in the prevention or treatment of cancer.
                4)- Blood pressure
                Maintaining a low sodium intake helps to keep blood pressure healthy; however, increasing potassiumintake may be just as important because of its vasodilation effects. According to the National Health and Nutrition Examination Survey, fewer than 2 percent of U.S. adults meet the daily 4700 mg recommendation.
                Also of note, a high potassium intake is associated with a 20 percent decreased risk of dying from all causes.
                 5)- Heart health
                The fiber, potassium, vitamin C and choline content in tomatoes all support heart health. An increase in potassium intake along with a decrease in sodium intake is the most important dietary change that the average person can make to reduce their risk of cardiovascular disease, according to Mark Houston, M.D., M.S., an associate clinical professor of medicine at Vanderbilt Medical School and director of the Hypertension Institute at St. Thomas Hospital in Tennessee.3 Tomatoes also contain folic acid, which helps to keep homocysteine levels in check, thereby reducing a risk factor for heart disease.
             In one study, those who consumed 4069 mg of potassium per day had a 49 percent lower risk of death from ischemic heart disease compared with those who consumed less potassium (about 1000 mg per day).
              High potassium intakes are also associated with a reduced risk of stroke, protection against loss of muscle mass, preservation of bone mineral density and reduction in the formation of kidney stones.
                6)- Diabetes
           Studies have shown that people with type 1 diabetes who consume high-fiber diets have lower blood glucose levels, while people with type 2 diabetes may have improved blood sugar, lipids and insulin levels. One cup of cherry tomatoes provides about 2 grams of fiber.
               The Dietary Guidelines for Americans recommends 21-25 g of fiber per day for women and 30-38 g/day for men.
                 7)- Constipation
               Eating foods that are high in water content and fiber like tomatoes can help with hydration and promote regular bowel movements. Fiber adds bulk to stool and is essential for minimizing constipation.
                 8)- Eye health
             Tomatoes are a rich source of lycopene, lutein and beta-carotene, powerful antioxidants that have been shown to protect the eyes against light-induced damage associated with the development of cataractsand age-related macular degeneration (AMD). The Age-Related Eye Disease Study (AREDS) recently found that people with high dietary intake of lutein and zeaxanthin (both carotenoids found in tomatoes) had a 35 percent reduction in the risk of neovascular AMD.
                9)- Skin
              The synthesis of collagen, an essential component of the skin, hair, nails and connective tissue, is reliant on vitamin C. A deficiency of vitamin C leads to scurvy. As vitamin C is a powerful antioxidant, a low intake is associated with increased damage from sunlight, pollution and smoke, leading to wrinkles, sagging skin, blemishes and other adverse health effects.
                10)- Pregnancy
                Adequate folic acid intake is essential before and during pregnancy to protect against neural tube defects in infants.
                 11)- Depression
              The folic acid in tomatoes may help with depression by preventing an excess of homocysteine from forming in the body, which can prevent blood and other nutrients from reaching the brain. Excess homocysteine interferes with the production of the feel-good hormones serotonin, dopamine, and norepinephrine, which regulate mood, sleep and appetite.
                Source: Tomatoes: Health Benefits, Facts, Research - Medical News Today

​                4.2- Varieties and Cultivars
                 + Varieties
               There are around 7,500 tomato varieties grown for various purposes having been selected with varying fruit types, and for optimum growth in differing growing conditions.
                 Tomato varieties can be divided into categories based on shape and size. Some important tomato varieties are listed below:
                 - Beefsteak tomatoes are 10 cm (4 in) or more in diameter, often used for  sandwiches and similar applications. Their kidney-bean shape, thinner skin, and shorter shelf life makes commercial use impractical.
                - Plum tomatoes, or paste tomatoes (including pear tomatoes), are bred with a lower water /higher solids content for use intomato sauce and paste, for canning and sauces and are usually oblong 7-9 cm (3-4 in) long and 4-5 cm (1.6-2.0 in) diameter; like the Roma-type tomatoes, important cultivars in the Sacramento Valley.
                 - Cherry tomatoes are small and round, often sweet tomatoes, about the same 1-2 cm (0.4-0.8 in) size as the wild tomato.
                 - Grape tomatoes are smaller and oblong, a variation on plum tomatoes.
               - Campari tomatoes are sweet and noted for their juiciness, low acidity, and lack of mealiness, bigger than cherry tomatoes, and smaller than plum tomatoes.
                 - Tomberries, tiny tomatoes, about 5 mm in diameter
                 - Oxheart tomatoes can range in size up to beefsteaks, and are shaped like large strawberries.
                 - Pear tomatoes are pear-shaped and can be based upon the San Marzano types for a richer gourmet paste.
                - "Slicing" or "globe" tomatoes are the usual tomatoes of commerce, used for a wide variety of processing and fresh eating. The most widely grown commercial tomatoes tend to be in the 5-6 cm (2.0-2.4 in) diameter range.
                 + Cutivars of the Species Solanum lycopersicum
                Some famous Cultivars of Tomatos are:
               1- Adoration
               2- Alicante
               3- Azoychka
               4- Beefsteak
               5- Better Boy
               6- Big Beef
               7- Big Rainbow
               8- Blaby Special
               9- Black Krim
               10- Brandywine
               11- Campari
               12- Celebrity
               13- Cherokee Purple
               14- Canario (tomato)
               15- Early Girl
               16- Enchantment
               17- Ferris Wheel
               18- Flamenco
               19- Fourth of July
               20- Garden Peach
               21- Gardener's Delight
               22- German Johnson
               23- Giulietta F1
               24- Granadero
               25- Great White
               26- Green Zebra
               27- Hillbilly
               28- Japanese Black Trifele
               29- Jersey Boy
               30- Jubilee
               31- Juliet
​32- Kumato
33- Lillian's Yellow
34- McDreamy
35- Matt's Wild Cherry
36- Micro Tom
37- Moneymaker
38- Monterosa
39- Montserrat
40- Mortgage Lifter
41- Mr. Stripey
42- Pantano Romanesco
43- Paul Robeson
44- Plum tomato
45- Pumpkin tomato
46- Raf tomato
47- Rebellion
48- Roma
49- Rosa de Barbastro (es)
50- Rutgers Tomato
51- San Marzano
52- Sasha Altai
53- Stupice
54- Tiny Tim
55- Traveller / Arkansas Traveler
56- Cherry Bambelo
57- Cherry Nebula
58- Santorini
59- Super Sweet 100
60- Tomaccio
61- Yellow Pear
62- White Queen
                                                                                                                                           Edited by Ho Dinh Hai                                                                                                                                                                                Long An - Vietnam
                                                                                       References
                1- http://www.worldwidefruits.com
                2- http://www.worldwidefruits.com/orders-of-plants.html
                3- https://en.wikipedia.org/wiki/Solanales
                4- http://www.mobot.org/mobot/research/apweb/orders/solanalesweb.htm
                5- https://en.wikipedia.org/wiki/Solanaceae
                6- https://en.wikipedia.org/wiki/Solanoideae
                7- https://en.wikipedia.org/wiki/Solanum
                8- https://en.wikipedia.org/wiki/Tomato                                                                                                      
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